Applejack is a hardworking previously Earth Pony from the town of Ponyville. The thread revolves around her ascension to alicorn status and her subsequent trials and tribulations as a princess, primarily tasked with filling out paperwork and righting wrongs from the Royal Sisters' long and passive rule.
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The white alicorn very likely swings back and forth on whether or not she is happy she made Applejack a princess. On the one hand, a lot of problems she had previously overlooked are being dealt with. On the other hand... A LOT of problems she overlooked need to be dealt with. While Celestia is proud of the strides made and all the good that's been done, she's also about two second from jumping off that balcony again. ONE MORE QUESTION AND LASERS, APPLEJACK!
When she was faced with a mountain of paperwork and more than anything needed a hand, who did she turn to? Why, one of the few characters who actually have hands, of course. Eager to continue being the number one assistant, and more than a little desperate to be noticed and feel like he could be heard, he went to work helping her out. Over time, they began to grow a lot closer. Maybe it was the confined space, maybe it was the reliance on each other, but something sparked, and now she's in a (comical) battle with Rarity for the dragons affection. It will probably end well... probably.
When Applejack decided to show the Bug mercy and try to help her out, it's very doubtful she invisioned her life being filled with sappy movies, chaotic assholes ruining her ballroom and the underlying scheming nature of the Changeling Queen hounding her at every turn. That said, do not be under the impression Applejack is going to take a single ounce of her attitude, and will toss a stapler in her face if she tries to slam her door. She's not exactly friends with Chrysalis, and likely never will be, but at the very least, they make for good allies if the need arises.
Do you really think the "War" for Spike's affection would ever make these two enemies? Of course not, Rarity is still her best friend through and through. Now that the fashionista has slimmed down, the pair are back to spending more time together and making Equestria a better place.
When she came up to the sad sack of sorry, she was having none of that. She realized she needed to be brutally honest with him in order to get him to stop acting like a neglected ninny and start being the Prince he needed to be. Her methods were... less than effective, at the start, but they did spark enough of a flame for him to get his own fire going, and now he's the one she can look to in the Crystal Empire for some sanity, besides 18 of course.
- Wait, why is she a princess again?
- Because she seems to be the only goddamn one of those ponies who gets that a Princess need to do actual work regarding maintenance of the kingdom. You know how much city planning or trade negotiations Twilight or Luna have done? Fucking nothing. Everyone is obsessed with the world ending threats they think I should fight, but the minute I point out the free healthcare I have to work to maintain everyone goes quiet. And don't even get me started on Cadence, who can't even manage a basic meeting with the Equestrian Games representative. Fucking annoying. Stop bothering her and let her go do actual princess stuff, because apparently everyone else got the pamphlets mixed up or something and thinks "Princess" means "Beat cop."
- She also has a rap, along with the two royal sisters. Because of course. https://soundcloud.com/applesnack/p